The energy used to pump hydrogen ions from the mitochondrial matrix across the inner membrane and into the intermembrane space is derived from

Potential energy is use to pump hydrogen ions or protons, from the inter-membrane space of the mitochondria into the matrix space. The energy is derived from the active pump that generates a proton concentration gradient across inner mitochondrial membrane then the difference in electric charge and pH creates an electrochemical potential, which produces potential energy.

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Single strand binding protein by peptide bond