In organisms with large genomes, inversions are more likely to be tolerated if the breakpoints occur in:

The answer is telomeres. These are the regions at the end of the chromosomes that are composed of random repeats of nucleotide sequences. When the DNA is duplicated entirely by DNA polymerases such as during mitosis, the lengths of chromosomes reduce slightly by every duplication. This is due to the aspect of the lagging strands replication that is unable to be completed to the end of the chromosome. Therefore telomeres protect the chromosome from losing critical genes since mutations on telomeres will not affect the cell functionality.  

parasites can detrimentally affect their host, but also must not allow their host to die before they can complete their life cycle. some parasites have complex life cycles with many different hosts, and have had to find ways of persisting in many different environments during the change from one host to another. ectoparasites live outside the body of the host, and can more easily  disperse, but are  more exposed to predators, parasites, parasitoids. endoparasites live inside the body of the host, and are therefore protected from the outside environment. but they can be  attacked by the host's immune system. they have therefore developed  mechanisms for dispersal such as  complex life cycles.

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