The yield of atp by the mitochondrial atp synthase can be estimated by the number of protons pumped out of the matrix during the oxidation of nadh and fadh 2 , and the number of protons moved into the matrix as atp is formed. mitochondrial oxidation of nadh by o 2 results in the transport of ten protons from the matrix to the intermembrane space, while oxidation of fadh 2 results in the transport of six protons. atp synthase utilizes three protons from the intermembrane space to form one atp . use the information given above to make the following calculations. note: recent data on atp synthase suggests that the number of protons required per atp varies with the number of c subunits. here, an approximate value of 3 protons / atp is used. how many atp are formed per nadh oxidized?